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=> résultent d’une perméabilité non contrôlée de la membrane plasmique :1) brèche2) hypoxie3) chute pdction ATP4) Na + H20 entre ds la cellule et le K sort de la cellule5) augmentation de la pression osmotique6) entré plus conséquente d'eau dans la cellule7) les citernes du RE se dilatent, rompent, et forment des vacuoles8) vacuolisation entensive9) dégénération hydropique
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