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One student was studying a null mutant X- in lac operon and thought there is one null mutation in the repressor gene (I), promoter (P), or operator (O). The student performed several experiments detecting the structural gene expression under medium with or without the inducer as well as with or without transformed F' with wild-type alleles (I+ P+ O+). Below is the data (-: no expression; +: expression). What is the most possible location of the null mutation?

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